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Adventure Press, National Geographic. Archived from the original on 31 December Bulls have a fibro-elastic penis. While the females do this, the penis is not withdrawn and research has shown a positive relationship between length of the time that the penis is licked and the duration of copulation. Handbook of the Birds of the World. Species of this genus have sex-reversed genitalia. Male echidnas have a four-headed penis.

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Archived from the original on 20 March When mating, the tip of a male pronghorn 's penis is often the first part to touch the female pronghorn. Functional Anatomy and Physiology of Domestic Animals. Cheetahs of the Serengeti Plains: Archived from the original on September 5, The male genitalia of mouse deer are similar to those of pigs. Male turtles and crocodiles have a penis, while male specimens of the reptile order Squamata have two paired organs called hemipenes.

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